Mottled Tortoise Beetle in Montgomery Co., Maryland (7/15/2024). (c) Jim Moore (Maryland), some rights reserved (CC BY-NC).
A Mottled Tortoise Beetle in Montgomery Co., Maryland (7/31/2018).
A Mottled Tortoise Beetle in Howard Co., Maryland (8/4/2014).
Mottled Tortoise Beetle in Wicomico Co., Maryland. Collected by Brooke Alexander and photographed by Amber Reese.
A Mottled Tortoise Beetle in Howard Co., Maryland (6/4/2018).
Mottled Tortoise Beetle in Montgomery Co., Maryland (5/8/2023). (c) Zachary Dankowicz, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Montgomery Co., Maryland (10/21/2020). (c) Katja Schulz, some rights reserved (CC BY).
Mottled Tortoise Beetle in Montgomery Co., Maryland (5/8/2023). (c) Zachary Dankowicz, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Wicomico Co., Maryland (5/31/2025). (c) Jim Moore (Maryland), some rights reserved (CC BY-NC).
A Mottled Tortoise Beetle in Howard Co., Maryland (6/22/2019).
A Mottled Tortoise Beetle in Montgomery Co., Maryland (8/16/2020).
Mottled Tortoise Beetle in Carroll Co., Maryland (7/17/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Montgomery Co., Maryland (6/6/2022). (c) David, some rights reserved (CC BY-NC-SA).
Mottled Tortoise Beetle in Montgomery Co., Maryland (6/6/2022). (c) David, some rights reserved (CC BY-NC-SA).
Mottled Tortoise Beetle in Montgomery Co., Maryland (6/2/2022). (c) belby, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Montgomery Co., Maryland (6/2/2022). (c) belby, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Montgomery Co., Maryland (7/26/2021). (c) Stephen John Davies, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Montgomery Co., Maryland (7/8/2021). (c) Stephen John Davies, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Montgomery Co., Maryland (5/5/2021). (c) maragay, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Prince George's Co., Maryland (7/10/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Prince George's Co., Maryland (7/10/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Prince George's Co., Maryland (7/13/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Prince George's Co., Maryland (6/3/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Prince George's Co., Maryland (6/3/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Prince George's Co., Maryland (6/20/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Calvert Co., Maryland (Date obscured). (c) Judy Ferris, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Calvert Co., Maryland (Date obscured). (c) Judy Ferris, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Calvert Co., Maryland (7/29/2021). (c) joshandleah, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Calvert Co., Maryland (7/29/2021). (c) joshandleah, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Anne Arundel Co., Maryland (7/31/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Anne Arundel Co., Maryland (7/31/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Anne Arundel Co., Maryland (6/13/2022). (c) fake swayze, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Anne Arundel Co., Maryland (5/22/2021). (c) adroege, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Baltimore Co., Maryland (6/5/2023). (c) rlkays, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Baltimore Co., Maryland (6/27/2023). (c) Frode Jacobsen, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Baltimore Co., Maryland (5/14/2023). (c) Douglas Hooper, some rights reserved (CC BY).
Mottled Tortoise Beetle in Baltimore Co., Maryland (6/5/2021). (c) Frode Jacobsen, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Baltimore Co., Maryland (10/3/2021). (c) nailles, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Worcester Co., Maryland (6/27/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
Mottled Tortoise Beetle in Dorchester Co., Maryland (9/10/2023). (c) Lisa Mayo, some rights reserved (CC BY-NC).
Mottled Tortoise Beetle in Washington Co., Maryland (7/17/2025). No rights reserved.
Mottled Tortoise Beetle in Montgomery Co. (9/14/2025). (c) Ryan Douglas, some rights reserved (CC BY-NC)
Map Snapshot
151 Records
Citations
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Mottled Tortoise Beetle in Montgomery Co., Maryland (7/15/2024). (c) Jim Moore (Maryland), some rights reserved (CC BY-NC).
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A Mottled Tortoise Beetle in Montgomery Co., Maryland (7/31/2018).
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A Mottled Tortoise Beetle in Howard Co., Maryland (8/4/2014).
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Mottled Tortoise Beetle in Wicomico Co., Maryland. Collected by Brooke Alexander and photographed by Amber Reese.
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A Mottled Tortoise Beetle in Howard Co., Maryland (6/4/2018).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (5/8/2023). (c) Zachary Dankowicz, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (10/21/2020). (c) Katja Schulz, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (5/8/2023). (c) Zachary Dankowicz, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Wicomico Co., Maryland (5/31/2025). (c) Jim Moore (Maryland), some rights reserved (CC BY-NC).
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A Mottled Tortoise Beetle in Howard Co., Maryland (6/22/2019).
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A Mottled Tortoise Beetle in Montgomery Co., Maryland (8/16/2020).
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Mottled Tortoise Beetle in Carroll Co., Maryland (7/17/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (6/6/2022). (c) David, some rights reserved (CC BY-NC-SA).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (6/6/2022). (c) David, some rights reserved (CC BY-NC-SA).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (6/2/2022). (c) belby, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (6/2/2022). (c) belby, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (7/26/2021). (c) Stephen John Davies, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (7/8/2021). (c) Stephen John Davies, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Montgomery Co., Maryland (5/5/2021). (c) maragay, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (7/10/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (7/10/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (7/13/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (6/3/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (6/3/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Prince George's Co., Maryland (6/20/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Calvert Co., Maryland (Date obscured). (c) Judy Ferris, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Calvert Co., Maryland (Date obscured). (c) Judy Ferris, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Calvert Co., Maryland (7/29/2021). (c) joshandleah, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Calvert Co., Maryland (7/29/2021). (c) joshandleah, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Anne Arundel Co., Maryland (7/31/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Anne Arundel Co., Maryland (7/31/2022). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Anne Arundel Co., Maryland (6/13/2022). (c) fake swayze, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Anne Arundel Co., Maryland (5/22/2021). (c) adroege, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Baltimore Co., Maryland (6/5/2023). (c) rlkays, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Baltimore Co., Maryland (6/27/2023). (c) Frode Jacobsen, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Baltimore Co., Maryland (5/14/2023). (c) Douglas Hooper, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Baltimore Co., Maryland (6/5/2021). (c) Frode Jacobsen, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Baltimore Co., Maryland (10/3/2021). (c) nailles, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Worcester Co., Maryland (6/27/2023). (c) Adrienne van den Beemt, some rights reserved (CC BY).
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Mottled Tortoise Beetle in Dorchester Co., Maryland (9/10/2023). (c) Lisa Mayo, some rights reserved (CC BY-NC).
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Mottled Tortoise Beetle in Washington Co., Maryland (7/17/2025). No rights reserved.
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Mottled Tortoise Beetle in Montgomery Co. (9/14/2025). (c) Ryan Douglas, some rights reserved (CC BY-NC)
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Source: Wikipedia
| Deloyala guttata | |
|---|---|
| |
Scientific classification 
| |
| Kingdom: | Animalia |
| Phylum: | Arthropoda |
| Clade: | Pancrustacea |
| Class: | Insecta |
| Order: | Coleoptera |
| Suborder: | Polyphaga |
| Infraorder: | Cucujiformia |
| Family: | Chrysomelidae |
| Genus: | Deloyala |
| Species: | D. guttata
|
| Binomial name | |
| Deloyala guttata (Olivier, 1790)
| |
Deloyala guttata, the mottled tortoise beetle, is a species of tortoise beetle in the family Chrysomelidae.[1][2] It is found in the Caribbean, Central America, North America, and South America.[2][3][4]
Larva carry shields made from exuviae and frass on their abdomen that they lift to shield their dorsum from predators.[6]
References
[edit]- ^ "Deloyala guttata Species Information". BugGuide.net. Retrieved 4 February 2018.
- ^ a b "Deloyala guttata Report". Integrated Taxonomic Information System. Retrieved 4 February 2018.
- ^ "North American Cryptocephalus species (Chrysomelidae, Cryptocephalinae)". Texas Entomology. Mike Quinn. Retrieved 4 February 2018.
- ^ "Staines C.L. (2011). "Catalog of the hispines of the World"". Smithsonian Institution. Retrieved 4 February 2018.
- ^ "Images and collection data for Deloyala guttata". Pictures from Earth. Retrieved 4 February 2018.
- ^ Olmstead, Karen L.; Denno, Robert F. (1993). "Effectiveness of Tortoise Beetle Larval Shields Against Different Predator Species". Ecology. 74 (5): 1394–1405. Bibcode:1993Ecol...74.1394O. doi:10.2307/1940069. ISSN 0012-9658. JSTOR 1940069. Retrieved 13 May 2022.
Further reading
[edit]- Arnett, R. H. Jr., M. C. Thomas, P. E. Skelley and J. H. Frank. (eds.). (21 June 2002). American Beetles, Volume II: Polyphaga: Scarabaeoidea through Curculionoidea. CRC Press LLC, Boca Raton, Florida ISBN 978-0-8493-0954-0.
- Arnett, Ross H. (2000). American Insects: A Handbook of the Insects of America North of Mexico. CRC Press.
- Borowiec, Lech (1999). A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae), 476.
- Richard E. White. (1983). Peterson Field Guides: Beetles. Houghton Mifflin Company.
- Riley, Edward G., Shawn M. Clark, and Terry N. Seeno (2003). "Catalog of the leaf beetles of America north of Mexico (Coleoptera: Megalopodidae, Orsodacnidae and Chrysomelidae, excluding Bruchinae)". Coleopterists Society Special Publication no. 1, 290.
- White, Richard E. (1968). A Review of the Genus Cryptocephalus in America North of Mexico. Smithsonian Institution Press.
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